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Rubus pensilvanicus

Poir. (1804)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Rosales Family: Rosaceae Genus: Rubus

Description

Key Characters:

Growth Form: Erect or becoming arching shrubs.

Stems: Stems stiff and usually erect in open areas, ± decumbent in shaded sites, primocanes 10–30 dm long, angled, covered with stout, straight or hooked prickles up to 6(–8) mm long, their bases usually longitudinally elongated, also sparsely pilose, lateral branches of floricanes moderately to densely pilose, prickles usually smaller.

Roots:

Leaves: Leaves compound (palmate). Alternate. Blades elliptic, oblong–oblanceolate, or ovate, leaflets (3)5, the terminal one usually 8–13 cm long, 3.5–5(–8) cm wide, terminal leaflet with longest petiolule usually 2–4 cm long, petiolules and petioles usually pilose and with prickles; floricane leaves similar to those of primocanes but smaller, leaflets 3(5), the terminal one usually only 5–9 cm long. Upper surfaces glabrate; lower surfaces usually soft pilose and midrib usually with small prickles. Margins coarsely and irregularly serrate. Petiolate. Stipules adnate to base of petioles.

Flowers: Flowers in short, open racemes, pedicels 15–50 mm long, moderately to densely villous (hairs not glandular or sometimes some of them glandular) and with prickles. Flowers bisexual (perfect), actinomorphic. Calyx lobes 5, persistent, distinct, imbricate. Corolla of 5 petals, white, distinct, obovate, imbricate, 13–20 mm long. Stamens numerous; filaments slender; anthers dithecal. Ovary superior, inserted on the raised receptacle, 1-celled; ovules 2 per cell; style filiform, subterminal, persistent.

Fruit: Fruit fleshy; juicy drupelets; black at maturity; subglobose to oblong; 1.5–2 cm long; cohering to the tardily separating receptacle. Seeds 1 per drupelet.

Ploidy: 2n = 14; 21

Habitat: Disturbed habitats; including mesic to wet forest and subalpine grassland.

Elevation Range: 200–3,000 m.

Historical Distribution

Synonyms (102)

Rubus abactus L.H.Bailey (1932)
Rubus ablatus L.H.Bailey (1945)
Rubus amnicola Blanch. (1906)
Rubus andrewsianus Blanch. (1906)
Rubus angustus L.H.Bailey (1945)
Rubus argutus var. scissus L.H.Bailey (1925)
Rubus arrectus L.H.Bailey (1945)
Rubus arvensis L.H.Bailey (1945)
Rubus associus Hanes ex L.H.Bailey (1947)
Rubus barbarus L.H.Bailey (1945)
Rubus bellobatus L.H.Bailey (1945)
Rubus blakei L.H.Bailey (1945)
Rubus bractealis L.H.Bailey (1947)
Rubus brainerdii Rydb. (1913)
Rubus burnhamii L.H.Bailey (1945)
Rubus bushii L.H.Bailey (1932)
Rubus cardianus L.H.Bailey (1945)
Rubus cauliflorus L.H.Bailey (1945)
Rubus condensiflorus L.H.Bailey (1945)
Rubus congruus L.H.Bailey (1945)
Rubus cupressorum Fernald (1947)
Rubus defectionis Fernald (1942)
Rubus demareanus L.H.Bailey (1945)
Rubus densipubens L.H.Bailey (1947)
Rubus densissimus H.A.Davis & T.Davis (1953)
Rubus difformis L.H.Bailey (1945)
Rubus eriensis L.H.Bailey (1948)
Rubus facetus L.H.Bailey (1932)
Rubus fatuus L.H.Bailey (1945)
Rubus frondosus (Torr.) Bigelow (1824)
Rubus fructifer L.H.Bailey (1945)
Rubus gattingeri L.H.Bailey (1945)
Rubus gnarus L.H.Bailey (1944)
Rubus harperi L.H.Bailey (1945)
Rubus heterogeneus L.H.Bailey (1945)
Rubus honorus L.H.Bailey (1934)
Rubus humilior L.H.Bailey (1947)
Rubus impar L.H.Bailey (1934)
Rubus independens L.H.Bailey (1947)
Rubus insons L.H.Bailey (1932)
Rubus insulanus L.H.Bailey (1945)
Rubus interioris L.H.Bailey (1949)
Rubus jennisonii L.H.Bailey (1945)
Rubus jugosus L.H.Bailey (1945)
Rubus kansanus L.H.Bailey (1945)
Rubus kelloggii L.H.Bailey (1945)
Rubus latifoliolus L.H.Bailey (1945)
Rubus laudatus A.Berger (1925)
Rubus leggii H.A.Davis & T.Davis (1953)
Rubus libratus L.H.Bailey (1945)
Rubus limulus L.H.Bailey (1945)
Rubus litoreus L.H.Bailey (1945)
Rubus missouricus L.H.Bailey (1932)
Rubus mollior L.H.Bailey (1945)
Rubus monongaliensis L.H.Bailey (1947)
Rubus multispinus Blanch. (1907)
Rubus nescius L.H.Bailey (1945)
Rubus offectus L.H.Bailey (1941)
Rubus oklahomus L.H.Bailey (1932)
Rubus oppositus L.H.Bailey (1949)
Rubus originalis L.H.Bailey (1945)
Rubus ozarkensis L.H.Bailey (1945)
Rubus paludivagus Fernald (1940)
Rubus pauxillus L.H.Bailey (1932)
Rubus pensilvanicus f. albinus E.J.Palmer & Steyerm. (1958)
Rubus pensilvanicus f. phyllophorus Fernald (1949)
Rubus pensilvanicus var. frondosus (Bigelow ex Torr.) B.Boivin (1967)
Rubus perfoliosus L.H.Bailey (1945)
Rubus perpauper L.H.Bailey (1945)
Rubus persistens Rydb. (1903)
Rubus philadelphicus Blanch. (1907)
Rubus praepes L.H.Bailey (1945)
Rubus pratensis L.H.Bailey (1945)
Rubus prestonensis H.A.Davis & T.Davis (1953)
Rubus pubifolius L.H.Bailey (1945)
Rubus pulchriflorus L.H.Bailey (1945)
Rubus putus L.H.Bailey (1945)
Rubus ramifer L.H.Bailey (1945)
Rubus recurvans Blanch. (1904)
Rubus recurvans var. subrecurvans Blanch. (1906)
Rubus rosarius L.H.Bailey (1925)
Rubus rydbergianus L.H.Bailey (1945)
Rubus saepescandens L.H.Bailey (1945)
Rubus schneckii L.H.Bailey (1945)
Rubus scibilis L.H.Bailey (1945)
Rubus senilis L.H.Bailey (1947)
Rubus sertatus L.H.Bailey (1945)
Rubus subsolanus L.H.Bailey (1945)
Rubus subtentus L.H.Bailey (1945)
Rubus subtractus L.H.Bailey (1945)
Rubus summotus L.H.Bailey (1945)
Rubus suus L.H.Bailey (1945)
Rubus texanus L.H.Bailey (1947)
Rubus tygartensis H.A.Davis & T.Davis (1953)
Rubus ucetanus L.H.Bailey (1925)
Rubus uniquus L.H.Bailey (1945)
Rubus villosus var. frondosus Bigelow ex Torr. (1824)
Rubus virilis L.H.Bailey (1945)
Rubus vixargutus L.H.Bailey (1945)
Rubus wahlii L.H.Bailey (1945)
Rubus wiegandii L.H.Bailey (1945)
Rubus zoae L.H.Bailey (1945)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents

Birds

Vegetative Reproduction

Pollinators

Specimens

Bibliography

Name Published In: J.B.A.M.de Lamarck, Encycl. 6: 246 (1804)

Other References

Wagner et al. 1990:1107 (K, O, M, H); Hughes 1995:9 (Mo)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Rubus pensilvanicus Poir. up Mauna Loa Strip Rd. ~0.5 mi from intersection with Kona-Hilo Hwy. (plot 18) Metrosideros, native shrub, scattered lichen on ash covered pahoehoe lava PRESERVED_SPECIMEN Newell, C.L. 798 Hawaii BISH 9/8/1966
2 Rubus pensilvanicus Poir. Mt. Kaala top, wet forest PRESERVED_SPECIMEN Christophersen, E. 1703 Oahu BISH 4/3/1931
3 Rubus pensilvanicus Poir. Kilauea, Military camp edges of thickets in disturbed ground, compact ash PRESERVED_SPECIMEN Fosberg, F.R. 33295 Hawaii BISH 2/12/1951
4 Rubus pensilvanicus Poir. East Maui, NE outer slope of Haleakala, N of Puu Alaea with Dubautia, Geranium, Sadleria, Vaccinium, Styphelia; in heath scrub PRESERVED_SPECIMEN Henrickson, J.S. 3673 Maui BISH 6/22/1969
5 Rubus pensilvanicus Poir. mt. kaala wet summit PRESERVED_SPECIMEN st.john, h. collector number: 22274 Oahu BISH 11/17/1946
6 Rubus pensilvanicus Poir. Kalaupulii, Kokee PRESERVED_SPECIMEN Degener, O. 21583 Kauai BISH 1/12/1952
7 Rubus pensilvanicus Poir. N.Kona, Holualoa Forest Reserve, S slope of Hualalai, near reservoir PRESERVED_SPECIMEN Kondo, Y. s.n. Hawaii BISH 7/13/1946
8 Rubus pensilvanicus Poir. ukulele, above olinda on steep windward facing slopes. Vegetation: Metrosideros, Myrsine, Ilex, Dodonaea, Wikstroemia, Leptecophylla, Chamaesyce, Lysimachia, Remya, Dryopteris, Alyxia, Microlepia, Carex. PRESERVED_SPECIMEN forbes, c.n. collector number: 166.m Maui BISH 7/1/1910
9 Rubus pensilvanicus Poir. kokee his garden PRESERVED_SPECIMEN lorence, d.h. flynn, t.; o'connor, p. collector number: 5752 Kauai BISH 1/28/1988
10 Rubus pensilvanicus Poir. Keauhou Ranch, Kau District Along fenceline of koa reforestation area. PRESERVED_SPECIMEN Cuddihy, L.W. ESP team Collector Number: 049 Hawaii BISH 12/18/1979
11 Rubus pensilvanicus Poir. up mauna loa strip rd. ~0.5 mi. from intersection with kona-hilo hwy. (plot 18) Metrosideros, native shrub, scattered lichen on ash covered pahoehoe lava PRESERVED_SPECIMEN newell, c.l. collector number: 798 Hawaii BISH 9/8/1966
12 Rubus pensilvanicus Poir. Kilauea, CCC camp, National Park PRESERVED_SPECIMEN Stone, B.C. 3105 Hawaii BISH 8/1/1959
13 Rubus pensilvanicus Poir. National Park, Kilauea, Residence #21 PRESERVED_SPECIMEN Fagerlund, G.O. 693 Hawaii BISH 7/30/1943
14 Rubus pensilvanicus Poir. n.kona, holualoa forest reserve, s slope of hualalai, near reservoir mesic forest, 60% closed 10-12 m canopy; Metrosideros, Acacia, Dodonaea viscosa, Santalum freycinetianum, Melicope ovata, M. barbigera, Poa sandwicensis, Gahnia beecheyi PRESERVED_SPECIMEN kondo, y. Kauai BISH 7/13/1946
15 Rubus pensilvanicus Poir. Olinda, Idlewild PRESERVED_SPECIMEN Krauss, N.L.H. 493 Maui BISH 3/19/1960
16 Rubus pensilvanicus Poir. east maui, koolau forest reserve, near puu alaea Dense ohia forest PRESERVED_SPECIMEN harrison, b. collector number: 495 Kauai BISH 10/10/1973
17 Rubus pensilvanicus Poir. olinda PRESERVED_SPECIMEN faurie, a. collector number: 801 Maui BISH 8/8/1909
18 Rubus pensilvanicus Poir. olinda Moist woods PRESERVED_SPECIMEN faurie, a. collector number: 801 Molokai BISH 8/8/1909
19 Rubus pensilvanicus Poir. Ukulele, above Olinda woods PRESERVED_SPECIMEN Forbes, C.N. 166.M Maui BISH 7/1/1910
20 Rubus pensilvanicus Poir. Waimea, Mahanaloa Valley, Kopiwai trail PRESERVED_SPECIMEN Doty, M.S. s.n. Kauai BISH 2/5/1952
21 Rubus pensilvanicus Poir. Halealoha PRESERVED_SPECIMEN Forbes, C.N. 797.H Hawaii BISH 6/10/1915
22 Rubus pensilvanicus Poir. on mauna loa strip road ~3.6 mi. from intersection with kona-hilo highway, into forest (plot 39) Acacia, Sapindus, Microlepia forest PRESERVED_SPECIMEN newell, c.l. collector number: 788 Hawaii BISH 9/8/1966
23 Rubus pensilvanicus Poir. Ukulele PRESERVED_SPECIMEN Forbes, C.N. 973.M Maui BISH 7/29/1919
24 Rubus pensilvanicus Poir. Kokee his garden PRESERVED_SPECIMEN Lorence, D.H. 5752 Kauai BISH 1/28/1988
25 Rubus pensilvanicus Poir. ukulele Metrosideros with Santalum, Dodonaea, Dicranopteris, Antidesma, Alphitonia, Psychotria, Pittosporum. PRESERVED_SPECIMEN forbes, c.n. collector number: 910.m Kauai BISH 7/24/1919
26 Rubus pensilvanicus Poir. ukulele PRESERVED_SPECIMEN forbes, c.n. collector number: 973.m Maui BISH 7/29/1919
27 Rubus pensilvanicus Poir. nw outer slope of haleakala, in hosmer's grove below national park Woods. PRESERVED_SPECIMEN henrickson, j.s. collector number: 3872 Hawaii BISH 7/15/1969
28 Rubus pensilvanicus Poir. Mt. Kaala PRESERVED_SPECIMEN St. John, H. 10064 Oahu BISH 11/24/1929
29 Rubus pensilvanicus Poir. down chain of craters rd. ~1.3 mi. past volcano observatory toward halemaumau crater parking lot (plot 51) Metrosideros, native shrub, scattered lichen on ash covered pahoehoe lava PRESERVED_SPECIMEN newell, c.l. collector number: 447 Hawaii BISH 8/19/1966
30 Rubus pensilvanicus Poir. east maui, olinda grazing land PRESERVED_SPECIMEN krauss, n.l.h. collector number: 614 Maui BISH 11/7/1961
31 Rubus pensilvanicus Poir. East Maui, Olinda, Kula Pipe Line PRESERVED_SPECIMEN St. John, H. 10311 Maui BISH 2/12/1930
32 Rubus pensilvanicus Poir. Mt. Kaala wet summit PRESERVED_SPECIMEN St. John, H. 22274 Oahu BISH 11/17/1946
33 Rubus pensilvanicus Poir. Thurston Lava Tube parking lot, ~3 m toward Tube, turn right up narrow jeep road (plot 4) Cibotium forest PRESERVED_SPECIMEN Newell, C.L. 930 Hawaii BISH 6/20/1967
34 Rubus pensilvanicus Poir. kilauea, ccc camp, national park PRESERVED_SPECIMEN stone, b.c. collector number: 3105 Oahu BISH 8/8/1959
35 Rubus pensilvanicus Poir. down Chain of Craters Rd. ~1.3 mi past Volcano Observatory toward Halemaumau Crater parking lot (plot 51) Metrosideros, native shrub, scattered lichen on ash covered pahoehoe lava PRESERVED_SPECIMEN Newell, C.L. 447 Hawaii BISH 8/19/1966
36 Rubus pensilvanicus Poir. keauhou ranch, kau district Along fenceline of koa reforestation area. PRESERVED_SPECIMEN cuddihy, l.w. esp team collector number: 049 Hawaii BISH 12/18/1979
37 Rubus pensilvanicus Poir. NW outer slope of Haleakala, in Hosmer's Grove below National Park along stream crossing road through grove; with conifers PRESERVED_SPECIMEN Henrickson, J.S. 3872 Maui BISH 7/15/1969
38 Rubus pensilvanicus Poir. Glenwood, 29 miles roadside PRESERVED_SPECIMEN Degener, O. 8179 Hawaii BISH 6/11/1929
39 Rubus pensilvanicus Poir. kilauea, military camp edges of thickets in disturbed ground, compact ash PRESERVED_SPECIMEN fosberg, f.r. collector number: 33295 Hawaii BISH 2/12/1951
40 Rubus pensilvanicus Poir. mt. kaala PRESERVED_SPECIMEN hosaka, e.y. collector number: 1267 Oahu BISH 10/4/1934
41 Rubus pensilvanicus Poir. Keauhou Ranch, Kau District Along fenceline of koa reforestation area. PRESERVED_SPECIMEN Cuddihy, L.W. 49 Hawaii BISH 12/18/1979
42 Rubus pensilvanicus Poir. kokee, puu kila lookout with Dicranopteris; Metrosideros rain forest, more open areas PRESERVED_SPECIMEN spence, g. collector number: 94 Kauai BISH 11/6/1972
43 Rubus pensilvanicus Poir. mt. kaala Acacia/Metrosideros lowland mesic forest with Dodonaea, Dianella, Gahnia, Styphelia PRESERVED_SPECIMEN st.john, h. collector number: 10064 Kauai BISH 11/24/1929
44 Rubus pensilvanicus Poir. glenwood, 29 miles roadside PRESERVED_SPECIMEN degener, o. collector number: 8179 Hawaii BISH 6/11/1929
45 Rubus pensilvanicus Poir. Big Island Hawai'i, Akaka Falls Lush areas beside parking lot. PRESERVED_SPECIMEN B.A. Bennett; R.S. Mulder 07-0326 Consortium of Pacific Northwest Herbaria 11/29/2007
46 Rubus pensilvanicus Poir. ukulele PRESERVED_SPECIMEN forbes, c.n. collector number: 910.m Maui BISH 7/24/1919
47 Rubus pensilvanicus Poir. mauna loa, e slopes above kilauea PRESERVED_SPECIMEN krajina, v.j. 25 HO 12/26/1961
48 Rubus pensilvanicus Poir. PRESERVED_SPECIMEN dina kageler FLAS 10/6/1982
49 Rubus pensilvanicus Poir. Mauna Loa, E slopes above Kilauea PRESERVED_SPECIMEN Krajina, V.J. 25 HO 12/26/1961
50 Rubus pensilvanicus Poir. West Maui Mts. PRESERVED_SPECIMEN Krauss, N. L. 1137 Maui US 4/27/1967
51 Rubus pensilvanicus Poir. Kalaupuhi, Kokee. PRESERVED_SPECIMEN Otto Degener 21583 Kauai PTBG 1/12/1952
52 Rubus pensilvanicus Poir. Kokee State Park, along Hwy 550 near mile 18.4. Site of proposed DOFAW's Kalalau Rim Exclosure. Metrosideros dominated wet forest. PRESERVED_SPECIMEN Mirutse Giday & Sara Friberg MGSF10 Kauai PTBG 7/23/1998
53 Rubus pensilvanicus Poir. Idlewild, Olinda PRESERVED_SPECIMEN Krauss, N. L. 493 Maui US 3/19/1960
54 Rubus pensilvanicus Poir. Keauhou Ranch. Along fencelina of Koa reforestation area. PRESERVED_SPECIMEN L. W. Cuddihy 49 Hawaii PTBG 12/18/1979
55 Rubus pensilvanicus Poir. Kawaiiki Valley, off Kaluahaulu ridge, N facing slope, below the ridge and just East of the smaller more eastern Diellia erecta f. alexandri fence. Mixed Mesic Forest PRESERVED_SPECIMEN Natalia Tangalin 1975 Kauai PTBG 4/9/2009
56 Rubus pensilvanicus Poir. Alakai Swamp, high elevation. Moist area. PRESERVED_SPECIMEN Helen Rubin s.n. Kauai PTBG 5/29/1975
57 Rubus pensilvanicus Poir. Kokee State Park, Waineke Swamp along Mohihi Road, ca. 0.5 miles E of Highway 550. marshy area with mixed mesophytic forest around perimeter. PRESERVED_SPECIMEN David H. Lorence 5752 Kauai PTBG 1/28/1988
58 Rubus pensilvanicus Poir. Hawaii Is., mauna loa, e. slopes above kilauea PRESERVED_SPECIMEN Vladimir J. Krajina 611226025 UBC 12/26/1961
59 Rubus pensilvanicus Poir. mauna kea, n side PRESERVED_SPECIMEN a. s. hitchcock 14297 Oahu US 8/22/1916
60 Rubus pensilvanicus Poir. kilauea military camp kilauea PRESERVED_SPECIMEN f. r. fosberg 33295 Hawaii US 2/12/1951
61 Rubus pensilvanicus Poir. kula pipleline w of waikamoi stream, haiku uka PRESERVED_SPECIMEN f. r. fosberg 41484 Maui US 3/14/1961
62 Rubus pensilvanicus Poir. Mt. Kaala, Waianae Mts. PRESERVED_SPECIMEN Nagata, K. M. 934 Oahu US 2/19/1972
63 Rubus pensilvanicus Poir. Olinda PRESERVED_SPECIMEN Krauss, N. L. 1034 Maui US 3/29/1966
64 Rubus pensilvanicus Poir. E Maui, Olinda, Kula Pipe Line PRESERVED_SPECIMEN St. John, H. 10311 Maui US 2/12/1930
65 Rubus pensilvanicus Poir. olinda, e of PRESERVED_SPECIMEN a. s. hitchcock 14933 Maui US 10/1/1916
66 Rubus pensilvanicus Poir. Mt. Kaala PRESERVED_SPECIMEN Davis, C. J. s.n. Oahu US 4/16/1966
67 Rubus pensilvanicus Poir. Haleakala Ranch pasture near Makawao For Res. PRESERVED_SPECIMEN Herbst, D. R.; Ishikawa, S. 4087 Maui US 8/27/1974
68 Rubus pensilvanicus Poir. Olinda PRESERVED_SPECIMEN Krauss, N. L. 614 Oahu US 1/7/1961
69 Rubus pensilvanicus Poir. koolau forest reserve, e maui, puu alaea Grassland PRESERVED_SPECIMEN b. harrison 495 Maui US 10/10/1973
70 Rubus pensilvanicus Poir. woods near ukulele, above olinda, e maui PRESERVED_SPECIMEN c. n. forbes 166.m. Maui US
71 Rubus pensilvanicus Poir. mt. kaala PRESERVED_SPECIMEN c. davis s.n. Oahu US 4/16/1966
72 Rubus pensilvanicus Poir. mt. kaala, waianae mts. PRESERVED_SPECIMEN k. nagata 934 Oahu US 2/19/1972
73 Rubus pensilvanicus Poir. waimea district, kokee state park, waineke swamp along mohihi road, ca. 0.5 mi e of highway 550 PRESERVED_SPECIMEN d. h. lorence, t. w. flynn & p. o'connor 5752 Kauai US 1/28/1988
74 Rubus pensilvanicus Poir. west maui mts. PRESERVED_SPECIMEN n. krauss 1137 Kauai US 4/27/1967
75 Rubus pensilvanicus Poir. e maui, olinda, kula pipe line PRESERVED_SPECIMEN h. st. john 10311 Maui US 2/12/1930
76 Rubus pensilvanicus Poir. along pipe-line trail, olinda PRESERVED_SPECIMEN a. tam 8800 Maui US
77 Rubus pensilvanicus Poir. Along Pipe-Line Trail, Olinda PRESERVED_SPECIMEN Tam, A. A. 8800 Maui US 10/1/1934
78 Rubus pensilvanicus Poir. idlewild, olinda PRESERVED_SPECIMEN n. krauss 493 Maui US 3/19/1960
79 Rubus pensilvanicus Poir. kaholuamano PRESERVED_SPECIMEN a. s. hitchcock 15425 Kauai US 10/21/1916
80 Rubus pensilvanicus Poir. Kilauea Military Camp Kilauea PRESERVED_SPECIMEN Fosberg, F. R. 33295 Hawaii US 2/12/1951
81 Rubus pensilvanicus Poir. kokee state park, around parking area at puu kila lookout PRESERVED_SPECIMEN g. spence 94 Kauai US 11/6/1972
82 Rubus pensilvanicus Poir. Koolau Forest Reserve, E Maui, Puu Alaea Grassland PRESERVED_SPECIMEN Harrison, B. 495 Maui US 1973-00-00
83 Rubus pensilvanicus Poir. olinda PRESERVED_SPECIMEN n. krauss 614 Oahu US 1/7/1961
84 Rubus pensilvanicus Poir. olinda PRESERVED_SPECIMEN n. krauss 1034 Maui US 3/29/1966
85 Rubus pensilvanicus Poir. Kokee State Park, around parking area at Puu Kila Lookout PRESERVED_SPECIMEN Spence, G. 94 Kauai US 11/6/1972
86 Rubus pensilvanicus Poir. Waimea District, Kokee State Park, Waineke Swamp along Mohihi Road, ca. 0.5 mi E of Highway 550 PRESERVED_SPECIMEN Lorence, D. H.; Flynn, T. W.; O'Connor, P. 5752 Kauai US 1/28/1988
87 Rubus pensilvanicus Poir. Mauna Kea, N side PRESERVED_SPECIMEN Hitchcock, A. S. 14297 Hawaii US 8/22/2016
88 Rubus pensilvanicus Poir. Olinda, E of PRESERVED_SPECIMEN Hitchcock, A. S. 14933 Maui US 10/1/2016
89 Rubus pensilvanicus Poir. Kaholuamano PRESERVED_SPECIMEN Hitchcock, A. S. 15425 Kauai US 10/21/2016
90 Rubus pensilvanicus Poir. Woods near Ukulele, above Olinda, E Maui PRESERVED_SPECIMEN Forbes, C. N. 166.M. Maui US 7/10/2020
91 Rubus pensilvanicus Poir. kokee state park. waialeale mountain range. PRESERVED_SPECIMEN s. starratt 389 WTU 3/27/1980
92 Rubus pensilvanicus Poir. Kula Pipleline W of Waikamoi Stream, Haiku Uka PRESERVED_SPECIMEN Fosberg, F. R. 41484 Maui US 3/14/1961
93 Rubus pensilvanicus Poir. Kokee, Puu Kila Lookout with Dicranopteris; Metrosideros rain forest, more open areas PRESERVED_SPECIMEN Spence, G. 94 Kauai BISH 11/6/1972
94 Rubus pensilvanicus Poir. near Glenwood, on the way to Volcano House PRESERVED_SPECIMEN Degener, O. 30188 Hawaii BISH 7/23/1926
95 Rubus pensilvanicus Poir. east maui, ne outer slope of haleakala, n of puu alaea PRESERVED_SPECIMEN henrickson, j.s. vogl, r. collector number: 3673 Kauai BISH 6/22/1969
96 Rubus pensilvanicus Poir. haleakala ranch near makawao forest reserve pasture PRESERVED_SPECIMEN herbst, d.r. ishikawa, s. collector number: 4037 Maui BISH 8/27/1974
97 Rubus pensilvanicus Poir. waianae mts., makaha valley Collected from the edge of the reef in the surf zone, depth approx. 15 ft. PRESERVED_SPECIMEN forbes, c.n. Oahu BISH 2/12/1909
98 Rubus pensilvanicus Poir. mt. kaala summit bog PRESERVED_SPECIMEN nagata, k.m. collector number: 934 Oahu BISH 2/19/1972
99 Rubus pensilvanicus Poir. east maui, koolau forest reserve, near puu alaea In tussock grassland with Deschampsia. PRESERVED_SPECIMEN harrison, b. collector number: 495 Maui BISH 10/10/1973
100 Rubus pensilvanicus Poir. Mt. Kaala summit bog PRESERVED_SPECIMEN Nagata, K.M. 934 Oahu BISH 2/19/1972
101 Rubus pensilvanicus Poir. mt. kaala PRESERVED_SPECIMEN townsend, m.m. Oahu BISH 1/12/1941
102 Rubus pensilvanicus Poir. Kokee, Kalalau lookout PRESERVED_SPECIMEN Krauss, N.L.H. 880 Kauai BISH 3/10/1963
103 Rubus pensilvanicus Poir. national park, kilauea PRESERVED_SPECIMEN swezey, o.h. wife collector number: degener 8606 Oahu BISH 7/28/1934
104 Rubus pensilvanicus Poir. W. Maui, Honokahua. Growing in yard around Haela'au Cabin, Kaulalewelewe. Disturbed Metrosideros/Dicranopteris wet forest. PRESERVED_SPECIMEN Hank Oppenheimer H50058 Maui PTBG 5/28/2000
105 Rubus pensilvanicus Poir. kalaupulii, kokee PRESERVED_SPECIMEN degener, o. greenwell, a. collector number: 21583 Kauai BISH 1/12/1952
106 Rubus pensilvanicus Poir. East Maui, Olinda grazing land PRESERVED_SPECIMEN Krauss, N.L.H. 614 Maui BISH 11/7/1961
107 Rubus pensilvanicus Poir. thurston lava tube parking lot, ~3 m toward tube, turn right up narrow jeep road (plot 4) Cibotium forest PRESERVED_SPECIMEN newell, c.l. collector number: 930 Hawaii BISH 6/20/1967
108 Rubus pensilvanicus Poir. east maui, hana district, haleakala, naholoku at paliku cabin Along the beach. In this area it forms large, sprawling colonies. Aalii, koli, kaliko, buffel grass and koa hoale are also common in the plant community. PRESERVED_SPECIMEN higashino, p.k. collector number: 853 Kauai BISH 6/20/1975
109 Rubus pensilvanicus Poir. waimea, mahanaloa valley, kopiwai trail PRESERVED_SPECIMEN doty, m.s. Kauai BISH 2/5/1952
110 Rubus pensilvanicus Poir. East Maui, Hana District, Haleakala, Naholoku at Paliku Cabin in 20' deep shaded bed of intermittent stream W of Ranger Cabin, with Dryopteris, Myrsine, Cheirodendron PRESERVED_SPECIMEN Higashino, P.K. 853 Maui BISH 6/20/1975
111 Rubus pensilvanicus Poir. national park, kilauea, residence #21 PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 693 Hawaii BISH 7/30/1943
112 Rubus pensilvanicus Poir. near Kilauea Crater hotel PRESERVED_SPECIMEN Skottsberg, C.J.F. 1123 Hawaii BISH 9/17/1922
113 Rubus pensilvanicus Poir. W Maui, Lahaina District, Kaulalewelewe PRESERVED_SPECIMEN Krauss, N.L.H. 1137 Maui BISH 4/27/1967
114 Rubus pensilvanicus Poir. mt. kaala top, wet forest PRESERVED_SPECIMEN christophersen, e. wilder, g.p.; hume, e.p. collector number: 1703 Oahu BISH 4/3/1931
115 Rubus pensilvanicus Poir. Mt. Kaala PRESERVED_SPECIMEN Hosaka, E.Y. 1267 Oahu BISH 10/4/1934
116 Rubus pensilvanicus Poir. mt. kaala Kaolite clay, PRESERVED_SPECIMEN st.john, h. collector number: 10064 Oahu BISH 11/24/1929
117 Rubus pensilvanicus Poir. Keauhou Ranch. Kau District Along fenceline of koa reforestation area. PRESERVED_SPECIMEN Cuddihy, L.W. State ESP team Collector Number: 049 Hawaii BISH 12/18/1979
118 Rubus pensilvanicus Poir. olinda PRESERVED_SPECIMEN krauss, n.l.h. collector number: 1034 Maui BISH 3/29/1966
119 Rubus pensilvanicus Poir. Mt. Kaala summit; prostrate in Metrosideros forest PRESERVED_SPECIMEN Grant, M.L. 7480 Oahu BISH 10/4/1934
120 Rubus pensilvanicus Poir. east maui, hana district, e end of haleakala crater, 20 yards s of paliku ranger cabin pasture; with Holcus lanatus PRESERVED_SPECIMEN henrickson, j.s. vogl, r. collector number: 3506 Maui BISH 6/16/1969
121 Rubus pensilvanicus Poir. mt. kaala summit; prostrate in Metrosideros forest PRESERVED_SPECIMEN grant, m.l. collector number: 7480 Oahu BISH 10/4/1934
122 Rubus pensilvanicus Poir. olinda, pipe line trail PRESERVED_SPECIMEN apo lam, a. collector number: degener 8800 Maui BISH 10/1/1934
123 Rubus pensilvanicus Poir. Olinda PRESERVED_SPECIMEN Faurie, A. 801 Maui BISH 8/1/1909
124 Rubus pensilvanicus Poir. Ukulele PRESERVED_SPECIMEN Forbes, C.N. 910.M Maui BISH 7/24/1919
125 Rubus pensilvanicus Poir. halealoha PRESERVED_SPECIMEN forbes, c.n. collector number: 797.h Hawaii BISH 6/10/1915
126 Rubus pensilvanicus Poir. kokee, kalalau lookout Scattered Metrosideros - native shrub - lichen community on ash-covered pahoehoe lava PRESERVED_SPECIMEN krauss, n.l.h. collector number: 880 Hawaii BISH 3/10/1963
127 Rubus pensilvanicus Poir. East Maui, Hana District, E end of Haleakala Crater, 20 yards S of Paliku Ranger Cabin pasture; with Holcus lanatus PRESERVED_SPECIMEN Henrickson, J.S. 3506 Maui BISH 6/16/1969
128 Rubus pensilvanicus Poir. keauhou ranch. kau district Along fenceline of koa reforestation area. PRESERVED_SPECIMEN cuddihy, l.w. state esp team collector number: 049 Hawaii BISH 12/18/1979
129 Rubus pensilvanicus Poir. on Mauna Loa Strip Road ~3.6 mi from intersection with Kona-Hilo Highway, into forest (plot 39) Acacia, Sapindus, Microlepia forest PRESERVED_SPECIMEN Newell, C.L. 788 Hawaii BISH 9/8/1966
130 Rubus pensilvanicus Poir. Mt. Kaala PRESERVED_SPECIMEN Townsend, M.M. s.n. Oahu BISH 1/12/1941
131 Rubus pensilvanicus Poir. Olinda PRESERVED_SPECIMEN Krauss, N.L.H. 1034 Maui BISH 3/29/1966
132 Rubus pensilvanicus Poir. National Park, Kilauea PRESERVED_SPECIMEN Swezey, O.H. Degener 8606 Hawaii BISH 7/28/1934
133 Rubus pensilvanicus Poir. near glenwood, on the way to volcano house PRESERVED_SPECIMEN degener, o. collector number: 30188 Hawaii BISH 7/23/1926
134 Rubus pensilvanicus Poir. waianae mts., kamaileunu ridge, sw ridge separating waianae kai and makaha valleys PRESERVED_SPECIMEN wagner, w.l. takeuchi, w.; obata, j.; r.yee collector number: 5091 Oahu BISH 12/11/1983
135 Rubus pensilvanicus Poir. East Maui, Koolau Forest Reserve, near Puu Alaea In tussock grassland with Deschampsia. PRESERVED_SPECIMEN Harrison, B. 495 Maui BISH 10/10/1973
136 Rubus pensilvanicus Poir. near kilauea crater hotel PRESERVED_SPECIMEN skottsberg, c.j.f. collector number: 1123 Hawaii BISH 9/17/1922
137 Rubus pensilvanicus Poir. olinda, idlewild PRESERVED_SPECIMEN krauss, n.l.h. collector number: 493 Maui BISH 3/19/1960
138 Rubus pensilvanicus Poir. Olinda, pipe line trail PRESERVED_SPECIMEN Apo Lam, A. Degener 8800 Maui BISH 10/1/1934
139 Rubus pensilvanicus Poir. Keauhou Ranch. Kau District Along fenceline of koa reforestation area. PRESERVED_SPECIMEN Cuddihy, L.W. 49 Hawaii BISH 12/18/1979
140 Rubus pensilvanicus Poir. east maui, olinda, kula pipe line PRESERVED_SPECIMEN st.john, h. collector number: 10311 Maui BISH 2/12/1930
141 Rubus pensilvanicus Poir. Haleakala Ranch near Makawao Forest Reserve pasture PRESERVED_SPECIMEN Herbst, D.R. 4037 Maui BISH 8/27/1974
142 Rubus pensilvanicus Poir. Waianae Mts., Kamaileunu Ridge, SW ridge separating Waianae Kai and Makaha Valleys PRESERVED_SPECIMEN Wagner, W.L. 5091 Oahu BISH 12/11/1983
143 Rubus pensilvanicus Poir. halealoha Sprawling into stream. PRESERVED_SPECIMEN forbes, c.n. collector number: 797.h Maui BISH 6/10/1915
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